24 amino acids

ExperimentalLongevity

Humanin

Also known as: HN, Mitochondrial-derived peptide

Molecular weight
2687.10 Da
Formula
C119H204N34O32S2
CAS
330936-69-1
Routes
3

Humanin is a 24-amino acid mitochondria-derived peptide (MDP) encoded by the 16S ribosomal RNA region of mitochondrial DNA. Discovered in 2001 during a screen for factors that protect neurons from Alzheimer's disease-related toxicity, humanin represents a paradigm shift in our understanding of mitochondrial genetics — demonstrating that mitochondrial DNA encodes not only structural/enzymatic proteins but also small bioactive peptides with systemic signaling functions. Humanin's primary biological roles include cytoprotection against cellular stress, metabolic regulation, and anti-inflammatory activity. It declines with age, correlating with increasing vulnerability to age-related diseases. Synthetic analogs (HNG — humanin with Gly→Ser substitution at position 14) show 1000-fold enhanced potency and are being developed for neuroprotective and cardiometabolic applications.

For educational and research purposes only
Last updated:Check the research sources

Section 01

What it's used for

Alzheimer's: Cell and Animal Research

In cell and animal studies, humanin protects brain cells from damage caused by amyloid-beta, reducing cell death, oxidative stress, and neuron connection loss. In rats, it eased amyloid-beta-related memory problems, but didn't clear plaque.

Animal
Clinical wording

In cell-culture and animal studies, humanin protects neurons from amyloid-beta toxicity, reducing neuronal death and downstream effects such as oxidative stress, tau hyperphosphorylation, and synaptic loss. In a rat model, humanin also attenuated amyloid-beta-induced cognitive deficits. No study was found showing that humanin reduces amyloid-beta plaque burden itself.

Heart and Blood Vessel Research

Research shows humanin reduces heart damage from blocked-then-restored blood flow, slows artery hardening, and improves blood-vessel-lining function. People with higher blood humanin levels tend to have lower cardiovascular disease risk.

AnimalIn vitroHuman
Clinical wording

Research shows humanin reduces cardiac ischemia/reperfusion injury, inhibits atherosclerosis, and improves endothelial function. Circulating humanin levels inversely correlate with cardiovascular disease risk.

Metabolic Health in Aging

Researchers observed that humanin improves how the body responds to insulin, reduces fatty liver buildup, and supports mitochondrial cell-energy function. It may help counter the metabolic decline linked to aging cell-energy systems.

AnimalLimited data
Clinical wording

Humanin improves insulin sensitivity, reduces hepatic steatosis, and enhances mitochondrial function. It may counteract metabolic decline associated with aging mitochondrial dysfunction.

Mitochondrial Signaling Research Tool

Humanin is a key research tool for understanding mitochondrial-derived peptides — a newly discovered class of signals sent from mitochondria, the cell's energy factories, back to the nucleus and other parts of the cell.

In vitroAnimal
Clinical wording

Humanin serves as a key research tool for understanding mitochondrial-derived peptide signaling — a new class of retrograde signals from mitochondria to the nucleus and other cellular compartments.

Section 02

Mechanism of Action

Mechanism 01

A tiny protein made by mitochondria

  • Researchers found it while screening for genes that rescue nerve cells from death.
  • Those cells had been damaged by Alzheimer's-linked genes and by amyloid-beta, the plaque protein.
  • It failed to rescue cells dying from two other genetic causes, so the effect is selective.
  • Its 24-unit sequence sits inside a gene of mitochondrial DNA, the cell's own power-plant genome.
Clinical wording

A short open reading frame inside the mitochondrial 16S rRNA gene

Humanin was found by functional expression screening for cDNAs that rescue neuronal cells from death caused by several familial Alzheimer's disease genes and by amyloid-beta. The clone encoded a 24-amino-acid polypeptide that was transcribed and then secreted into the culture medium, and the rescue depended on its primary structure, with no effect on death caused by Q79 or SOD1 mutants. The identical sequence is present in mitochondrial DNA, inside the MT-RNR2 16S rRNA gene, which places humanin in the mitochondrial-derived peptide family.

Mechanism 02

Blocking the cell's self-destruct proteins

  • It grabs a protein called Bax and stops it reaching mitochondria to start cell death.
  • Silencing the peptide made cells more vulnerable and let more Bax move to membranes.
  • It also binds one long form of a related killer protein, but not the shorter forms.
  • All of these results come from cell lines and isolated mitochondria, not from animals.
Clinical wording

Bax and BimEL sequestration at the outer mitochondrial membrane

Humanin binds Bax and prevents its translocation from cytosol to mitochondria; reducing humanin with siRNA sensitises cells to Bax and increases Bax movement to membranes. Humanin also binds the extra-long isoform BimEL, but not BimL or BimS, and protects against BimEL-induced apoptosis, while humanin mutants unable to bind BimEL lose that protection. On mitochondria isolated from bax-null cells humanin still blocked BimEL-driven SMAC and cytochrome c release and prevented Bak oligomerisation. Cell lines and isolated organelles.

Mechanism 03

A three-part receptor on nerve cells

  • In two nerve cell lines the peptide made three receptor pieces clump together.
  • Treatment switched on STAT3, a messenger carrying survival signals towards the nucleus.
  • Silencing two of the receptor pieces cut the protective effect to roughly a fifth.
  • Injected into mice, it raised survival signalling in the hippocampus of old but not young animals.
Clinical wording

CNTFR-alpha/WSX-1/gp130 receptor complex and STAT3

In F11 neurohybrid and SH-SY5Y cells, humanin induced hetero-oligomerisation of the recombinant extracellular domains of CNTFR-alpha, WSX-1 and gp130, and immunoprecipitation showed humanin-dependent association of endogenous WSX-1 with gp130. Treatment raised STAT3 Tyr705 phosphorylation, and siRNA against CNTFR-alpha or WSX-1 cut humanin and HNG cytoprotection to roughly 20% of control. Acting through GP130/IL6ST, humanin also raises AKT, ERK1/2 and STAT3 phosphorylation via PI3K, MEK and JAK respectively, and injected humanin raised hippocampal AKT and ERK1/2 phosphorylation in old but not young mice.

Mechanism 04

Competing with the Alzheimer's plaque protein

  • It attracts immune scavenger cells through a receptor also used by amyloid-beta 42.
  • In nerve precursor cells both molecules switched that receptor on, but only amyloid-beta killed cells.
  • The peptide blocked that killing, consistent with the two competing for the same receptor.
  • Later high-resolution structures showed exactly where each molecule sits inside the receptor.
Clinical wording

FPR2/FPRL1, a receptor shared with amyloid-beta 42

Synthetic humanin induced chemotaxis of mononuclear phagocytes through human FPRL1 and its mouse counterpart FPR2, the same receptors amyloid-beta 42 uses. In neuroblast cells both peptides activated FPRL1, but only amyloid-beta 42 caused apoptotic death, and humanin blocked it, consistent with competition for receptor access. Cryo-EM structures of FPR2 bound to Gi with either amyloid-beta 42 or N-formyl humanin later localised recognition to a polar cavity inside the receptor helical bundle and a hydrophobic groove in the extracellular region.

Mechanism 05

Effects on blood sugar in rodents

  • A screening method identified a carrier protein (IGFBP-3) that binds the peptide directly.
  • Injected into rat brains, it lowered sugar output by the liver and raised uptake elsewhere.
  • That effect ran through a brain messenger, and blocking the messenger abolished it.
  • In mice with interrupted heart blood flow, a modified version raised two protective signals.
Clinical wording

IGFBP-3 binding and metabolic signalling in rodents

A yeast two-hybrid screen identified humanin as an IGFBP-3 binding partner; binding maps to the 18-amino-acid heparin-binding domain of IGFBP-3, does not compete with IGF-I, and is lost in the F6A variant. In rats, intracerebroventricular humanin during a hyperinsulinaemic-euglycaemic clamp lowered hepatic glucose production and raised peripheral glucose uptake, with an approximately tenfold rise in hypothalamic STAT3 phosphorylation that STAT3 inhibition abolished; the non-IGFBP-3-binding analogue was more potent. In mouse myocardial ischaemia-reperfusion the analogue HNG raised AMPK and eNOS phosphorylation.

Section 03

Biological Pathways

  1. STAT3 via tripartite receptorHumanin drives hetero-oligomerization of CNTFR-alpha, WSX-1 and gp130, raising STAT3 Tyr705 phosphorylation; silencing CNTFR-alpha or WSX-1 cut cytoprotection to roughly 20% of control.
  2. Bax and BimEL sequestrationHumanin binds Bax, preventing its translocation to mitochondria, and separately binds BimEL to block BimEL-induced apoptosis, still blocking cytochrome c release in bax-null cell lines and isolated organelles.
  3. FPR2/FPRL1 shared with amyloid-betaHumanin activates FPRL1/FPR2, the same receptors used by amyloid-beta 42, driving phagocyte chemotaxis; in neuroblast cells amyloid-beta 42 caused apoptosis through this receptor, and humanin blocked it.
  4. IGFBP-3 binding and AMPK/eNOSHumanin binds the heparin-binding domain of IGFBP-3 without competing with IGF-I; in rats this lowered hepatic glucose output via hypothalamic STAT3, while HNG raised AMPK and eNOS phosphorylation.

Section 04

Dosage Information

Amino acid sequence
MAPRGFSCLLLLTSEIDLPVKRRA
Ranges reported in experimental work
Route / systemContextRange studiedLimitation
Endogenous — measured, not administeredObservational human studies; nobody was given itMedian 774 pg/mL in sedentary people vs 1258 pg/mL in athletes; 2.2 ± 1.5 ng/mL with healthy heart vessels vs 1.3 ± 1.1 ng/mL withoutLevels the body keeps on its own and nobody changed, only compared with fitness and vessel health. They say nothing about injecting milligrams.
Into the abdomen — mouse studiesOld mice; HNG, a stronger lab-made versionHNG 4 mg/kg twice weekly from 18 months of age for 14 months; heart and metabolism models used 0.1–4 mg/kg a dayHNG is a strengthened humanin, injected into the belly of mice. 4 mg/kg would be 280–360 mg in a 70–90 kg adult — never scaled, measured or given to anyone.
Subcutaneous — self-administrationResearch-chemical vendors and community practice1–5 mg a day, or about 1 mg per session three times weekly in 4–12 week cycles — roughly 11–71 µg/kg for a 70–90 kg adultVendors say outright the numbers are scaled from mouse mg/kg. No registered trial has ever given humanin or HNG to a person, so nothing has tested them.
Dosage calculatorMass · concentration · volume · U-100

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Dosage calculation parameters

Substance mass shown on the vial label.

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70

The dose is derived from this weight and the mcg/kg rate.

mcg/kg

Research range 1.43–71.4 mcg/kg.

Calculated dose70 kg × mcg/kg

Results

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ConcentrationSubstance mass in one millilitre of solution.
Doses per VialComplete calculated doses, rounded down.

Set positive values for: Recommended dose per kg.

Research Use Only. This information is for educational and research purposes only. Not intended for medical advice or self-medication.

Section 05

Protocols

No protocols featuring this peptide yet. Browse All Protocols

Section 06

Stability & Storage

  1. Lyophilised powder

    Stored at −20 °C and shielded from air and light. The methionine at position 1 makes the peptide prone to oxidation, and the exact retention time depends on the batch and manufacturer.

  2. After reconstitution

    Reconstituted with sterile water shortly before use. Because the peptide oxidises easily, the solution is kept cold and used promptly, with the exact holding time following the manufacturer's instructions.

Section 07

Side Effects & Precautions

Limited human safety data as humanin remains largely in preclinical research. No significant toxicity observed in animal studies at therapeutic doses. Theoretical concern that anti-apoptotic activity could promote survival of damaged or pre-malignant cells.

Section 08

Regulatory Status

Humanin is not approved by any drug regulator in the world for human therapeutic use.

It remains a preclinical research peptide: no approved label, indication, dose or established human safety profile exists for it.

  1. FDA / United States

    Not approved

    Humanin has never been nominated for the 503A bulk-substances process and is not on the 503A Bulks List, so section 503A opens no lawful route for a pharmacy to compound it for human use.

  2. EU / EMA

    Not approved

    Humanin has no EU marketing authorisation and does not appear among medicines evaluated by the CHMP.

  3. WADA

    Prohibited under S0

    Humanin is not individually named on the Prohibited List, but as a mitochondria-derived peptide outside the growth-hormone/IGF-1 pathway covered by S2, it falls under the S0 catch-all for non-approved substances — banned at all times, with no therapeutic-use exemption.

  4. Clinical trials

    No completed human trial exists

    Evidence for humanin's cytoprotective and metabolic effects comes from cell cultures and rodent models; no Phase 1 safety or pharmacokinetic study in humans has been published.

A «research use only» label does not turn humanin into an approved medicine and says nothing about its purity. Regulatory status differs between jurisdictions and changes over time — check the current documents of your own regulator before relying on any of this.

Section 09

Research Studies

  1. [1]A rescue factor abolishing neuronal cell death by a wide spectrum of familial Alzheimer's disease genes and AbetaHashimoto Y, Niikura T, Tajima H, et al. · Proceedings of the National Academy of Sciences · 2001
  2. [2]Humanin peptide suppresses apoptosis by interfering with Bax activationGuo B, Zhai D, Cabezas E, et al. · Nature · 2003
  3. [3]Cytoprotective peptide humanin binds and inhibits proapoptotic Bcl-2/Bax family protein BimELLuciano F, Zhai D, Zhu X, et al. · Journal of Biological Chemistry · 2005
  4. [4]Humanin inhibits neuronal cell death by interacting with a cytokine receptor complex or complexes involving CNTF receptor alpha/WSX-1/gp130Hashimoto Y, Kurita M, Aiso S, Nishimoto I, Matsuoka M. · Molecular Biology of the Cell · 2009
  5. [5]The mitochondrial-derived peptide humanin activates the ERK1/2, AKT, and STAT3 signaling pathways and has age-dependent signaling differences in the hippocampusKim SJ, Guerrero N, Wassef G, et al. · Oncotarget · 2016
  6. [6]Humanin, a newly identified neuroprotective factor, uses the G protein-coupled formylpeptide receptor-like-1 as a functional receptorYing G, Iribarren P, Zhou Y, et al. · The Journal of Immunology · 2004
  7. [7]Structural basis of FPR2 in recognition of Abeta42 and neuroprotection by humaninZhu Y, Lin X, Zong X, et al. · Nature Communications · 2022
  8. [8]Interaction between the Alzheimer's survival peptide humanin and insulin-like growth factor-binding protein 3 regulates cell survival and apoptosisIkonen M, Liu B, Hashimoto Y, et al. · Proceedings of the National Academy of Sciences · 2003
  9. [9]Humanin: a novel central regulator of peripheral insulin actionMuzumdar RH, Huffman DM, Atzmon G, et al. · PLoS ONE · 2009
  10. [10]Acute humanin therapy attenuates myocardial ischemia and reperfusion injury in miceMuzumdar RH, Huffman DM, Calvert JW, et al. · Arteriosclerosis, Thrombosis, and Vascular Biology · 2010
  11. [11]Peptides derived from small mitochondrial open reading frames: genomic, biological, and therapeutic implicationsMiller B, Kim SJ, Kumagai H, et al. · Experimental Cell Research · 2020

Section 10

Frequently Asked Questions

Humanin is a 24-amino-acid peptide encoded inside mitochondrial DNA — inside the 16S ribosomal RNA gene, not a standard protein-coding gene — discovered in 2001 while screening for factors that protect neurons from Alzheimer's-related toxicity. In cell and animal studies it blocks programmed cell death by binding pro-apoptotic proteins like Bax and BimEL, and activates survival signalling (STAT3, AKT, ERK) through cell-surface receptors including a CNTFR-alpha/WSX-1/gp130 complex and FPR2 — the same receptor amyloid-beta 42 uses. It is classed as a mitochondrial-derived peptide, a signalling role for mitochondrial DNA identified only in the past two decades.

No. The only human data are observational: circulating humanin levels were measured and compared between groups — for example, a median of 774 pg/mL in sedentary people versus 1258 pg/mL in athletes, and higher levels in people with healthy coronary vessels than those without. Nobody was given humanin in these studies; everything about administering it, including the stronger synthetic analog HNG, comes from mouse experiments.

The only interventional dosing is in mice: old mice received the enhanced analog HNG at 4 mg/kg injected into the abdomen twice weekly for 14 months, with other mouse metabolic and cardiac studies using 0.1–4 mg/kg a day. Self-administration practice circulating outside any trial — roughly 1–5 mg a day — is a direct scale-up of that mouse mg/kg figure, something vendors state outright; no human dose-finding study of humanin or HNG has ever been conducted.

Animal studies at the doses tested showed no significant toxicity, but human safety data barely exist because humanin has never gone through a human trial. There is a specific theoretical concern rather than a measured one: humanin's core mechanism is blocking programmed cell death, and the same anti-apoptotic action that protects healthy neurons could in principle help damaged or pre-malignant cells survive. This has not been tested either way.

Humanin has no regulatory approval anywhere and remains an investigational compound in preclinical and early translational research. It is not on WADA's prohibited list, unlike many growth-factor peptides, since it has no established performance-enhancing use — that absence of prohibition reflects its research status, not a safety clearance.

As lyophilised powder it is kept at −20 °C, shielded from air and light — the methionine at position 1 makes it prone to oxidation, and how long it holds up varies by batch and manufacturer. Once reconstituted, the solution is kept cold and used promptly rather than stored, again because of that oxidation sensitivity.